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Types of Adaptations Relevant to Population Dynamics

Adoption of alternative reproductive tactics is another life-history adaptation to strong reproductive competition. Examples include adoption of sneaking versus territorial aggressive mating tactics in several fish species, for example, the blueguill sunfish (Lepomis macrochirus) whereby small and early maturing males become 'sneakers' that dart-in and ejaculate over the eggs of a copulating pair of large individuals. This alternative 'sneaking' life history (early maturing, small body size) is thought to increase

Pattern of sperm storage

During copulation the male constructs the spermatophore within the bursa copulatrix. Sperm transfer usually occurs towards the end of copulation after spermatophore construction (Callahan and Cascio, 1963). Some time after copulation, sperm leave the bursa copulatrix via the ductus seminalis and are transported, probably aided by female reproductive tract movements (Tschudi-Rein and Benz, 1990 LaMunyon and Eisner, 1993), to the spermatheca (Fig. 3.3). This means that there is little scope for males to directly interfere with sperm already stored from previous matings. Females store sperm in the spermatheca until required for fertilization, which occurs at oviposition. Eupyrene sperm leave the spermatheca through the spermathecal duct, and fertilization occurs when the egg passes the spermathecal duct opening as it travels from the oviduct to the ovipositor (Norris, 1932 Callahan and Cascio, 1963). There is therefore scope for ejaculates of more than one male to be stored in the...

Changing the dynamics of malefemale sexual conflict through alteration of the population sex ratio

It is clear that as well as overt mating system differences, these parasites are likely to produce more covert changes in the reproductive biology of the species concerned. In particular, we would expect male ejaculate constitution to change with increasing population sex ratio bias, and corresponding changes in the ability of males to mate multiply. Preliminary evidence for this comes from studies of the nymphalid butterfly Hypolimnas bolina (Dyson and Hurst, 2004) (see Fig. 8.1). This species inhabits the island and mainland areas of the Pacific, and the prevalence of a male-killing Wolbachia varies between islands. In one island, Independent Samoa, prevalence is extreme (99 of females are infected), and the population sex ratio is 100 females per male. On this island, unmatedness of females is common, indicating that male access to females has become unlimited. The size of spermatophore delivered by males during copulation in Independent Samoa is less than 50 of that found on...

Male compounds that may affect female immunity

Male insects are known to transfer compounds in their seminal fluids that affect female receptivity (e.g. Thornhill and Alcock, 1983), oviposition rates, and longevity (e.g. Chapman et al., 1995). Given that these physiologically active compounds affect a complex range of female traits it is likely that they may coincidently, or deliberately, enhance reduce female immune function. For example, it has been established that male Drosophila transfer three different antimicrobial peptides in their seminal fluid (Lung et al., 2001) as well as transfer compounds that activate phenoloxidase in the female's genital tract (Asada and Kitagawa, 1988). These male-transferred compounds may function primarily to protect and or enhance the competitiveness of the male's ejaculate, but may have the correlated effect of protecting the female and or reducing her immune costs as well. We know almost nothing about the immuno-logical function of insect seminal fluid and or why males transfer these...

Reproduction And Development

Each ejaculation releases 2 to 5 mL of semen, the fluid containing the sperm. Semen normally contains between 20 and 100 million sperm per milliliter. The sperm consists of a 5-p.m head containing the nucleus, a midpiece containing mitochondria, and a 55-p.m-long flagellum, or tail, which can propel the sperm at a speed of 1 to 4 mm min. The seminal vesicles and the prostate gland produce much of the liquid that makes up semen, which includes fructose to provide energy for the sperm, and alkalinity to counter the acidity of the vagina. When the male becomes sexually excited, nerves of the autonomic nervous system produce dilation of arterioles in the penis, causing blood to enter that organ faster than it can leave. This causes the erectile tissue to become engorged, causing an erection. The male then inserts the penis into the female's vagina in the act of copulation, injecting the semen by rhythmic contractions called ejaculations.

Passenger microorganisms and the design of spermatogenesis and oogenesis

As well as selection to accommodate passenger microorganisms within oogenesis spermatogenesis, there may also be selection to prevent their access to the germline. In the case of sex-ratio-distorting microorganisms, selection favours modifiers that prevent the microbe entering the ovary, as this prevents the progeny suffering the parasitic phenotype. In the case of Wolbachia inducing cytoplasmic incompatibility, selection may favour modifiers that prevent access of the Wolbachia to testes. This is for two reasons. First, it may prevent the cost of incompatibility. More importantly, it may prevent the cost to sperm production of having testes densely packed with Wolbachia. Snook et al. (2000) observed that D. simulans males infected with Wolbachia strain wRi had a lower rate of spermatogenesis, and fertilized fewer eggs per ejaculate.

Sperm number

Theory predicts that an increased number of sperm is advantageous in sperm competition, either because a larger ejaculate volume displaces more rival sperm or due to numerical superiority. In general, sperm competition favouring numerous sperm may have led to the evolution of many tiny sperm and may even be responsible for the evolution of the two sexes (Parker et al., 1972 Bulmer and Parker, 2002). Comparative data in numerous animal groups have corroborated this finding. Males invest more resources in testes and sperm production when they encounter a greater risk of sperm competition (Smith, 1984 Birkhead and M0ller, 1998 Simmons, 2001). The same is found in butterflies. In species where females mate more frequently, as determined by spermatophore counts in wild-caught females, males have relatively larger testes (Gage, 1994). As yet, we do not know whether these males also produce more sperm. However, it seems likely, as in other insects, that there is generally a positive...

Sperm Removal

At the conclusion of a successful copulation in cockroaches the transferred sperm are housed within a sper-matophore in the female genital tract. The male is long gone before his gametes move to the female spermathe-cae, and is likely to have little direct influence on where, how, and if his sperm are stored. If his female consort is not a virgin, however, there is potential for a copulating male to increase his fertilization success by using genital appendages to move or remove the stored sperm of a rival. Male intromittent organs are known to extract stored sperm in one of three basic ways (Eberhard, 1996 Miller, 1990). First, a genital structure may be inserted into or near a spermatheca and the ejaculate issued with enough force to flush out a rival's sperm. This mechanism is unlikely in cockroaches since sperm transfer is indirect, via a spermatophore. Second, male genital appendages may be used to induce the female to discard the sperm of other

Seminal factors

Substances transferred in the spermatophore play a role in switching off female receptivity and stimulating oviposition and egg maturation rate (Gillott, 2003). In the moth Heliothis virescens, female egg maturation is stimulated by juvenile hormone derived from the males' accessory glands (Park and Ramaswamy, 1998). In addition, it appears that male-derived factors also stimulate the female's own production of juvenile hormone (Park et al., 1998). In Helicoverpa zea and H. armigera other factors from the male accessory gland stimulate egg maturation and oviposition (Bali et al., 1996 Jin and Gong, 2001). In many moth species, receptive females attract males by releasing pheromones during a characteristic phase of 'calling' behaviour. The regulation of female pheromone production has been intensively studied in moths. A neuropeptide (PBAN) regulates pheromone production (e.g. Raina, 1993 Rafaeli, 2002), mediated by either humoral, hormonal or neural cues. There is large interspecific...

Sperm length

Accordingly, numerical sperm competition explains why there is selection for large numbers of sperm. Males are driven to minimize sperm size to maximize sperm number. On this basis, one might predict across related species which vary in mating pattern, that as sperm competition risk increases then sperm size should either decrease or remain minimal, both of which will enable sperm number to increase as investment in relative testis size increases. Across fish (Stockley et a ., 1997) this prediction is satisfied as sperm competition risk increases, testis size and ejaculate sperm number increase while sperm size decreases. However, across butterflies the opposite is found. Although relative testis size (Gage, 1994) and spermatophore mass (Svard and Wiklund, 1989) increase with degree of polyandry, there is also a positive increase in relative eupyrene sperm length with level of sperm competition (Gage, 1994). Fertile sperm are longer than would be expected in species which generate...

Stlv Stlv

Sun, 11 Apr 2021 | Infectious Diseases

Intromissions prior to ejaculation (Dewsbury and Pierce 1989 Dixson 1998), because this could increase the risk of micro-injury (abrasions, cuts) to the genitals and the total contact time for each copulation. Finally aggressive interactions among males that are competing for access to females could lead to the spread of disease (Tutin 2000).

Undiminished passion

The second was a serendipitous choice - the fowl Gallusgallus. In the late 1980s I was invited to a meeting at the University of Stockholm's field station at Tovetorp in southern Sweden. During a tour of the facilities we were shown enclosures containing lynx Lynx lynx, moose Alces alces and other macho large mammals, all of which were being studied by rather macho research students. Suddenly a group of feral fowl (a primitive domestic fowl very similar to the red jungle fowl) scuttled past us and a male forced a copulation almost at our feet. Taken aback, I asked my host which of the various research students was studying these birds. Slightly incredulous, he said 'no one' - they were simply 'decoration'. I was intrigued, and a year or two later Tom Pizzari (see Chapter 10) was there as my PhD student studying their behaviour. I had mentioned to Tom that if he could persuade the cockerels to copulate with a stuffed female we might be able to obtain natural ejaculates, as I had done...

Mate Choice

However, the most studied component of multiple mating concerns sperm competition. This widespread phenomenon occurs when sperm from two or more males compete for a female's ova. Relative sperm numbers are important for sperm competitive success, so that species experiencing higher rates of sperm competition have males with larger testes that produce more sperm. Sperm competition is therefore a selective force shaping optimal ejaculate structure, although spermato-genesis is not unlimited. While males usually possess a greater reproductive potential than females, males have evolved mechanisms for the optimal allocation of finite sperm among females to maximize lifetime reproductive success.

Sperm Competition

When the probability of female remating is high, selection should favor adaptations in males that allow them to reduce or avoid competition with the sperm of another male. This can lead to rapid and divergent evolution of traits that function in sperm competition and its avoidance. These traits may be manifest in behavior (e.g., mate guarding), genital morphology (e.g., structures that deliver sperm closer to the spermatheca), and physiology (e.g., chemicals in the ejaculate that enhance the success of sperm). Selection may also act at the level of the sperm itself, in that some may be adapted to outcompete others for access to eggs (Ridley, 1988 Eberhard, 1996 Simmons, 2001). Variation in Ejaculates A number of studies indicate that males increase the size of their ejaculate in the presence of rival males (summarized in Wedell et al., 2002). Harris and Moore (2004) tested the idea in N. cinerea by exposing adult males during their post-emergence maturation period to the chemical...

Spermathecal Shape

Two basic spermathecal shapes are represented in cockroaches the tubular form, with little difference in width between the duct and the spermatheca proper ( ampulla), and the capitate form, shaped like a lollipop. Shape varies widely across cockroach species and sometimes within a species. In Agmoblatta thaxteri each spermatheca has a double terminal bulb, like a figure 8 (Gurney and Roth, 1966). The genus Tryonicus can be inter- and in-traspecifically polymorphic (Fig. 6.18) (Roth, 1987b) however, some apparent variation in spermathecal shape may be due to the amount of ejaculate stored or to the preservation of specimens at different stages of muscular activity. Both the ampulla and ducts are surrounded by a sheath of profusely innervated striated muscle (Gupta and Smith, 1969). The sheath is best developed at the base, where it consists mainly of circular fibers and functions as a sphincter in opening and closing the entry (van Wyk, 1952). dict sperm use patterns (Walker, 1980),but...